Insects

Homoptera Classification

Several alternative classifications are in use for the Hemiptera-Heteroptera-Homoptera group of the Insecta. The classification adopted in Nomina Insecta Nearctica is to avoid the term Hemiptera and divide the complex into two orders Heteroptera and Homoptera. A commonly accepted alternative is to use a single order Hemiptera divided (in North America) into three suborders; Heteroptera, Sternorrhyncha, and Auchenorrhyncha. The Sternorrhyncha and Auchenorrhyncha are the two main subdivisions of the Homoptera in the sense of this check list. This latter classification seems to be gaining in acceptance.

The primary source for most of the Auchenorrhyncha is a monumental series of world catalogues compiled and published over a number of years by Z. P. Metcalf and generally entitled "A General Catalogue of the Homoptera".

CICADIDAE
Duffels, J.P., and P.A. Van Der Laan. 1985. Catalogue of the Cicadoidea (Homoptera,
Auchenorhyncha) 1956-1980. Dr. W. Junk Publishers. Dordrecht. 414 pp.
Cicadinae: Beameria, Cacama, Cicada, Cornuplura, Diceroprocta, Fidicina, Neocicada, Odopoea, Pacarina, Quesada, Zammara.
Platypediinae: Neoplatypedia, Platypedia.
Tettigadinae: Tettigades.
Tibicininae: Cicadetta, Clidophleps, Herrera, Magicicada, Melampsalta, Okanagana, Okanagodes, Tibicen, Tibicinoides.

CERCOPIDAE
The North American species of the Cercopidae are sometimes divided into three families corresponding to the subfamilies as listed here. Following the conservative path the Cercoidae are treated as a single family.
Aphrophorinae: Aphrophora, Lepyronia, Neophilaenus, Paraphilaenus, Philaenarcys, Philaenus, Philaronia.
Cercopinae: Prosapia.
Clastopterinae: Clastoptera.

AETALIONIDAE
Aetalion, Microcentrus.

MEMBRACIDAE
The Membracidae are based on; Metcalf, Z.P., and V. Wade. 1965. General Catalogue of the Homoptera. A Supplement to Fascicle I - Membracidae of the General Catalogue of the Hemiptera. In Two Volumes. Baltimore. Waverly Press. Section 1, pp. 1-585, Section 2, pp. 587-919.
Centrotinae: Centrodontus, Dontonodus, Gargara, Tylocentrus, Umbonia.
Hoplophorioninae: Platycotis.
Darniinae: Stictopelta.
Smiliinae: Acutalis, Amastris, Anisostylus, Antianthe, Aphetea, Archasia, Atymna, Bajulata, Bryantopsis, Carynota, Ceresa, Cyrtolobus, Entylia, Evashmeadea, Glossonotus, Grandolobus, Hadrophallus, Heliria, Helonica, Idioderma, Ilithucia, Micrutalis, Ophiderma, Palonica, Parantonae, Platycentrus, Polyglypta, Publilia, Smilia, Spissistilus, Stictocephala, Stictolobus, Telamona, Telamonanthe, Telonaca, Thelia, Tortistilus, Trachytalis, Trichaetipyga, Tropidarnis, Vanduzea, Vestistilus, Xantholobus.
Membracinae: Bolbonota, Campylenchia, Enchenopa, Hypsoprora, Leioscyta, Membracis, Multareis, Multareoides, Philya, Scalmophorus, Tylopelta.

CICADELLIDAE
Oman, P.W., Knight, W.J., and M.W. Nielson. 1990. Leafhoppers (Cicadellidae): A Bibliography, Generic Check-List and Index to the World Literature 1956-1985. CAB International. Wallingford. 368 pp.
Agalliinae: Aceratagallia, Agalliopsis, Agalliota, Ceratagallia.
Aphrodinae: Anoscopus, Aphrodes, Planaphrodes, Stroggylocephalus.
Cicadelliinae: Allogonia, Amblyscarta, Amphigonalia, Ankosus, Apogonalia, Aulacizes, Bathysmatophorus, Carsonus, Cicadella, Ciminius, Cuerna, Decua, Draeculacephala, Errhomus, Evacanthus, Friscanus, Graphocephala, Helochara, Homalodisca, Hortensia, Lystridea, Manzutus, Metascarta, Oncometopia, Pagaronia, Paraulacizes, Phera, Plesiommata, Sibovia, Thatuna, Tylozygus, Xyphon.
Coelidiinae: Harasupia, Jikradia, Tinobregmus.
Deltocephalinae: Acinopterus, Aflexia, Alapus, Allygianus, Allygiella, Allygus, Amblysellus, Amplicephalus, Aplanus, Aplanusiella, Arundanus, Ascius, Atanus, Athysanella, Athysanus, Auridius, Balclutha, Baldulus, Ballana, Bandara, Bonneyana, Boreotettix, Caladonus, Calanana, Calonia, Cantura, Cazenus, Cetexa, Chlorotettix, Cicadula, Cochlorhinus, Colladonus, Commellus, Cosmotettix, Coulinus, Crassana, Cribrus, Crumbana, Dalbulus, Daltonia, Danbara, Davisonia, Deltanus, Deltazotus, Deltella, Deltocephalus, Destria, Dicyphonia, Diplocolenus, Dixianus, Doleranus, Doliotettix, Doratura, Dorydiella, Drionia, Driotura, Elymana, Endria, Errastunus, Eulonus, Eusama, Euscelis, Eusora, Eutettix, Exitianus, Extrusanus, Fieberiella, Fitchana, Flexamia, Flexarida, Floridonus, Fridonus, Frigartus, Gillettiella, Giprus, Gloridonus, Graminella, Graphocraerus, Haldorus, Hardya, Hebecephalus, Hebexa, Hecalus, Hecullus, Huleria, Idiodonus, Japananus, Kansendria, Kinonia, Knullana, Laevicephalus, Latalus, Lemellus, Limbanus, Limotettix, Lonatura, Lycioides, Macrosteles, Memnonia, Mendozellus, Menosoma, Mesamia, Mexara, Mocuellus, Neoaliturus, Neodonus, Nephotettix, Nigridonus, Norvellina, Nurenus, Ollarianus, Omanana, Ophiola, Opsius, Orientus, Orocastus, Osbornellus, Paluda, Parabolocratus, Paramesus, Paraphlepsius, Pasadenus, Pasaremus, Pazu, Peconus, Pendarus, Penehuleria, Phlepsanus, Phlepsius, Pinumius, Planicephalus, Polyamia, Prescottia, Psammotettix, Reventazonia, Rosenus, Sanctanus, Scaphoideus, Scaphytopius, Sobara, Sorhoanus, Spangbergiella, Spartopyge, Spathanus, Stenometopiellus, Stirellus, Telusus, Texananus, Thamnotettix, Tideltellus, Tropicanus, Twiningia, Unoka, Zabrosa.
Eupelicinae: Attenuipyga, Dorycara, Neoslossonia.
Gyponinae: Acuera, Acusana, Dragonana, Gypona, Gyponana, Hamana, Marganana, Negosiana, Polana, Ponana, Prairiana, Rugosana, Scaris.
Iassinae: Momoria, Pachyopsis, Penestragania, Stragania.
Idiocerinae: Balcanocerus, Idiocerus, Jamacerus, Rhytidodus.
Koebeliinae: Koebelia.
Ledrinae: Xerophloea.
Macropsinae: Macropsis, Oncopsis, Pediopsis, Pediopsoides, Reticopsis.
Megophthalminae: Brenda, Tiaja.
Neocoelidiinae: Neocoelidia, Neocoelidiana.
Nioniinae: Nionia.
Penthimiinae: Penthimia.
Typhlocybinae: Aguriahana, Alconeura, Alebra, Chlorita, Diceratalebra, Dikraneura, Dikrella, Empoasca, Erythroneura, Eupterella, Eupteryx, Eurhadina, Fagocyba, Forcipata, Henribautia, Hymetta, Idona, Joruma, Kidrella, Kunzeana, Kunzella, Mcateeana, Notus, Ossiannilssonola, Protalebrella, Rabela, Ribautiana, Solanasca, Trypanalebra, Typhlocyba, Typhlocybella, Zygina.
Xestocephalinae: Xestocephalus.

DELPHACIDAE
Acanthodelphax, Achorotile, Araeopus, Aschedelphax, Bakerella, Bostaera, Chilodelphax, Chloriona, Copicerus, Criomorphus, Delphacodes, Dicranotropis, Euidella, Eurysa, Harmalia, Isodelphax, Javesella, Kelisia, Kelisoidea, Keyflana, Kormus, Kusnezoviella, Laccocera, Liburnia, Liburniella, Macrotomella, Megamelanus, Megamelus, Neomegamelanus, Nothodelphax, Paradelphacodes, Pareuidella, Parkana, Penepissonotus, Pentagramma, Peregrinus, Phyllodinus, Pissonotus, Prokelisia, Ribautodelphax, Rotundifronta, Saccharosydne, Sogata, Stenocranus, Stobaera, Tumidagena, Unkanodes, Yukonodelphax.

CIXIIDAE
Asotocixius, Bothriocera, Cixius, Microledrida, Monorachis, Myndus, Nymphocixia, Oecleus, Oliaronus, Oliarus, Pintalia, Platycixius, Stegocixius.

FULGORIDAE
Alphina, Amycle, Calyptoproctus, Cyrpoptus, Poblicia, Rhabdocephala, Scolopsella.

ACHILIDAE
Catonia, Epiptera, Juniperthia, Momar, Opsiplanon, Synecdoche, Uniptera, Xerbus.

DERBIDAE
Amalopota, Anotia, Apache, Cedusa, Cenchrea, Dysimia, Euklastus, Mula, Mysidia, Neocenchrea, Otiocerus, Patara, Persis, Phaciocephalus, Sayiana, Shellenius, Syntames.

DICTYOPHARIDAE
Acinaca, Aridia, Deserta, Loxophora, Mitrops, Nersia, Orgamara, Orgerius, Parorgerius, Phylloscelis, Rhynchomitra, Scolops, Ticida, Timodema, Timonidia, Yucanda.

TROPIDUCHIDAE
Monopsis, Neurotmeta, Pelitropis, Tangia.

KINNARIDAE
Oeclidius.

ISSIDAE
Acanalonia, Aphelonema, Asarcopus, Bruchomorpha, Caliscelis, Danepteryx, Dictydea, Dictyobia, Dictyonia, Dictyonissus, Dictyssa, Dictyssonia, Dyctidea, Euthiscia, Fitchiella, Hysteropterum, Issus, Misodema, Neaethus, Osbornia, Papagona, Picumna, Thionia, Traxus, Tylanira, Ulixes.

FLATIDAE
Anormenis, Antillormenis, Cyarda, Flatoides, Flatoidinus, Hesperophantia, Leptormenis, Melormenis, Metcalfa, Mistharnophantia, Ormenaria, Ormenis, Ormenoides, Pseudoflatoides.

The primary source of data for the Psylloidea is: Hodkinson, 1988, The Nearctic Psylloidea (Insecta: Homoptera): an annotated check list. Journal of Natural History, 22:1179-1243. The family and subfamily classification of the Psylloidea does not seem to be generally agreed upon by current workers. Therefore the family classification as given in the above reference is used, but with families arranged alphabetically.

APHALARIDAE
Aphalara, Aphalaroida, Blastopsylla, Craspedolepta, Ctenarytaina, Gyropsylla, Katacephala, Leurolophus, Limataphalara, Livia, Neophyllura, Paraphalaroida, Psyllopsis, Vailakiella.

CALOPHYIDAE
Calophya.

HOMOTOMIDAE
Homotoma.

PSYLLIDAE
Acizzia, Amorphicola, Arytaina, Arytainilla, Cacopsylla, Ceanothia, Euglyptoneura, Euphalerus, Heteropsylla, Nyctiphalerus, Pachypsylla, Pexopsylla, Psylla, Purshivora, Spanioneura, Tetragonocephala.

TRIOZIDAE
Ceropsylla, Choricymoza, Hemitrioza, Kuwayama, Leuronota, Levidea, Metatrioza, Neotriozella, Paratrioza, Rhinopsylla, Trioza.

ALEYRODIDAE
Aleurodicinae: Aleurodicus, Paraleyrodes.
Aleyrodinae: Aleurocanthus, Aleurochiton, Aleurocybotus, Aleuroglandulus, Aleurolobus, Aleuroparadoxus, Aleuroplatus, Aleuropleurocelus, Aleurothrixus, Aleurotithius, Aleurotulus, Aleyrodes, Bemisia, Crenidorsum, Dialeurodes, Paraleurolobus, Pealius, Tetraleurodes, Trialeurodes.

APHIDIDAE
The main source for data entry is: Smith, C.F., and C.S. Parron. 1978. An Annotated List of Aphididae (Homoptera) of North America. North Carolina Agricultural Experiment Station Technical Bulletin no. 255. 428 pp. The classification of the Aphididae is based on; Fottit, R.G. and W.R. Richards. 1993. The Insects and Arachnids of Canada. Part 22. The Genera of the Aphids of Canada. Homoptera: Aphidoidea and Phylloxeroidea. Agriculture Canada. Ottawa. 766 pp. The family Aphididae is split into a number of families in this publication. Each of the families in the above publication is treated as a subfamily in this classification to retain the conservative treatment of the Aphididae. Not all North American genera have been placed to subfamily in this classification.
Mindarinae: Mindarus.
Hormaphidinae: Atarsaphis, Cerataphis, Hamamelistes, Hormaphis.
Phloeomyzinae: Phloeomyzus.
Thelaxinae: Glyphina, Thelaxes.
Anoeciidae: Anoecia.
Pemphiginae: Colopha, Cornaphis, Eriosoma, Forda, Geoica, Geopemphigus, Grylloprociphilus, Kaltenbachiella, Melaphis, Meliarhizophagus, Mordvilkoja, Neoprociphilus, Pachypappa, Pemphigus, Prociphilus, Smynthurodes, Stagona, Tetraneura, Thecabius, Tiliphagus.
Drepanosiphinae: Appendiseta, Atheroides, Betulaphis, Boernerina, Callaphis, Callipterinella, Cepegillettea, Chaitophorus, Chromaphis, Ctenocallis, Diphyllaphis, Drepanaphis, Drepanosiphum, Eucallipterus, Euceraphis, Hoplochaitophorus, Iziphya, Lachnochaitophorus, Monellia, Monelliopsis, Myzocallis, Neosymydobius, Oestlundiella, Periphyllus, Phyllaphis, Protopterocallis, Pterocallis, Saltusaphis, Sipha, Stegophylla, Subiziphya, Subsaltusaphis, Symydobius, Takecallis, Tamalia, Therioaphis, Thripsaphis, Tinocallis, Tuberculatus.
Aphidinae: Acuticauda, Acyrthosiphon, Alphitoaphis, Amphicercidus, Amphorophora, Aphis, Aphthargelia, Artemisaphis, Asiphonaphis, Aspidaphis, Aspidaphium, Atarsos, Aulacorthum, Bipersona, Brachycaudus, Brachycolus, Brevicoryne, Brachycorynella, Brachyunguis, Braggia, Cachryphora, Calaphis, Capitophorus, Carolinaia, Catamergus, Cavariella, Cedoaphis, Ceruraphis, Chaetosiphon, Coloradoa, Cryptaphis, Cryptomyzus, Decorosiphon, Diuraphis, Dysaphis, Elatobium, Eoessigia, Eomacrosiphon, Epameibaphis, Ericaphis, Fimbriaphis, Flabellomicrosiphum, Fullawaya, Glendenningia, Gypsoaphis, Hayhurstia, Hyadaphis, Hyalomyzus, Hyalopteroides, Hyalopterus, Hyperomyzus, Hysteroneura, Idiopterus, Illinoia, Jacksonia, Landisaphis, Linosiphon, Liosomaphis, Lipamyzodes, Lipaphis, Longicaudus, Macrosiphoniella, Macrosiphum, Mastopoda, Megouroparsus, Melanaphis, Microlophium, Microparsus, Microsiphoniella, Misturaphis, Muscaphis, Myzaphis, Myzodium, Myzus, Nasonovia, Nearctaphis, Neoamphorophora, Neotoxoptera, Obtusicauda, Ovatus, Paducia, Papulaphis, Phorodon, Placoaphis, Pleotrichophorus, Pseudacaudella, Pseudocercidis, Pseudoepameibaphis, Pseudopterocomma, Pseudosiphonaphis, Pterocomma, Rhodobium, Rhopalomyzus, Rhopalosiphoninus, Rhopalosiphum, Sanbornia, Schizaphis, Siphonatrophia, Subacyrtosiphon, Toxoptera, Uroleucon, Utamphorophora, Vesiculaphis, Wahlgreniella, Xerophilaphis, Zyxaphis.
Lachninae: Cinara, Eulachnus, Essigella, Lachnus, Longistigma, Maculolachnus, Schizolachnus, Trama, Tuberolachnus.
Not placed to subfamily in database: Aloephagus, Anthracosiphon, Anuraphis, Brachysiphum, Byrsocryptoides, Clydesmithia, Delphiniobium, Deurocapillata, Euthoracaphis, Iowana, Macchatiella, Megoura, Megourina, Melanocallis, Nevadaphis, Paracolopha, Patchia, Pentalonia, Pentamyzus, Picturaphis, Pseudoregma, Sarucallis, Schizoneurata, Schoutedenia, Semiaphis, Titanosiphon.

PHYLLOXERIDAE
Pergande, T. 1904. North American Phylloxerinae affecting Hicoria (Carya) and other trees. Proceedings of the Davenport Academy of Sciences, 9:185-273.
Daktulosphaira, Phylloxera.

ADELGIDAE
Adelges, Pineus.

MARGARODIDAE
Crypticerya, Desmococcus, Dimargarodes, Heteromargarodes, Icerya, Kuwania, Margarodes, Matsucoccus, Neosteingelia, Pityococcus, Porphyrophora, Promargarodes, Steatococcus, Xyloccoculus.

ORTHEZIIDAE
Arctorthezia, Newsteadia, Nipponorthezia, Orthezia.

TACHARDIIDAE
Tachardiella.

COCCIDAE
Ben-Dov, Y. 1993. A Systematic Catalogue of the Soft Scale Insects of the World (Homoptera: Coccoidea: Coccidae) with Data on Geographical Distribution, Host Plants and Economic Importance. Sandhill Crane Press, Inc. Gainesville. 536 pp.
Ceroplastes, Ceroplastodes, Coccus, Eriopeltis, Eucalymnatus, Eulecanium, Exaeretopus, Inglisia, Kilifia, Lecanopsis, Luzulaspis, Mesolecanium, Metapulvinaria, Milviscutulus, Neolecanium, Neopulvinaria, Parasaissetia, Parthenolecanium, Philephedra, Physokermes, Protopulvinaria, Pseudokermes, Pseudophilippia, Pulvinaria, Saissetia, Toumeyella, Vinsonia.

ACLERDIDAE
Aclerda.

KERMESIDAE
Allokermes, Eriokermes, Kermes, Nanokermes, Olliffiella.

ASTEROLECANIIDAE
Asterolecanium, Mycetococcus, Pollinia, Sclerosococcus.

LECANODIASPIDIDAE
Lecanodiaspis.

CEROCOCCIDAE
Cerococcus.

DACTYLOPIIDAE
Dactylopius.

DIASPIDIDAE
Primary data entry from: Borchsenius, N.S. 1966. A Catalogue of the Armoured Scale Insects (Diaspidididae) of the World. Moscow. Nauka. 449 pp. and Nakahara, S. 1982. Checklist of the armored scales (Homoptera: Diaspididae) of the conterminous United States. 110 pp.
Abgrallaspis, Acutaspis, Ancepaspis, Andaspis, Annulaspis, Aonidia, Aonidiella, Aonidomytilus, Aspidaspis, Aspidiella, Aspidiotus, Aulacaspis, Carulaspis, Chionaspis, Chortinaspis, Chrysomphalus, Circulaspis, Clavaspis, Comstockiella, Crassaspis, Crenulaspidiotus, Cupidaspis, Dactylaspis, Diaspidiotus, Diaspis, Dicirculaspis, Duplaspidiotus, Dynaspidiotus, Epidiaspis, Exuviaspis, Ferrisidea, Fiorinia, Fissuraspis, Furcaspis, Furchadaspis, Genaparlatoria, Geodiaspis, Gymnaspis, Haliaspis, Hemiberlesia, Hemigymnaspis, Howardia, Ischnaspis, Kuwanaspis, Lapazia, Lepidosaphes, Leucaspis, Lindingaspis, Lopholeucaspis, Melanaspis, Morganella, Mycetaspis, Nelaspis, Neopinnaspis, Nicholiella, Niveaspis, Nuculaspis, Odonaspis, Opuntiaspis, Palinaspis, Pallulaspis, Paracupidaspis, Parlatoreopsis, Parlatoria, Pelliculaspis, Pinnaspis, Poliaspis, Praecocaspis, Protodiaspis, Pseudaonidia, Pseudaulacaspis, Pseudischnaspis,Pseudodiaspis, Pseudoparlatoria, Quadraspidiotus, Quernaspis, Radionaspis, Rhizaspidiotus, Rugaspidiotus, Selenaspidus, Situlaspis, Stramenaspis, Targionia, Unaspis, Velataspis, Xeropilaspis.

CONCHASPIDIDAE
Asceloconchaspis, Conchaspis.

PHOENICOCOCCIDAE
Phoenicococcus.

PSEUDOCOCCIDAE
This family is based on: Ben-Dov, Y. 1994. A Systematic Catalogue of the Mealybugs of the World (Insecta: Homoptera: Coccoidea: Pseudococcidae and Putoidae) with Data on Geographical Distribution, Host Plants, Biology and Economic Importance. Intercept Ltd. Andover. 686 pp. Although the author includes Putoidae in the title of the work, he does not accept the family concept in the body of the text.
Amonostherium, Anisococcus, Anthelococcus, Antonina, Antoninoides, Balanococcus, Brevennia, Cataenococcus, Chnaurococcus, Chorizococcus, Circaputo, Crisicoccus, Cryptoripersia, Cucullococcus, Discococcus, Distichlicoccus, Dysmicoccus, Ehrhornia, Eurycoccus, Ferrisia, Geococcus, Heliococcus, Heterococcus, Humococcus, Hypogeococcus, Misericoccus, Nipaecoccus, Oracella, Paludicoccus, Paracoccus, Paradoxococcus, Peliococcus, Phenacoccus, Planococcus, Plotococcus, Pseudantonina, Pseudococcus, Puto, Pygmaeococcus, Radicoccus, Rhizoecus, Ripersia, Scaptococcus, Spilococcus, Stemmatomerinx, Syrmococcus, Tridiscus, Trionymus, Vryburgia.

ERIOCOCCIDAE
Hoy, J.M. 1963. A Catalogue of the Eriococcidae (Homoptera: Coccoidea) of the World. New Zealand Department of Scientific and Industrial Research. Bulletin 150. 260 pp.
Acanthococcus, Apezococcus, Atriplicia, Cornoculus, Cryptococcus, Gossyparia, Hypericicoccus, Oregomopyga, Ovaticoccus, Spiroporococcus.